Intense solar radiation constrains plant species richness in global grasslands

Authors

Marie Spohn, Sveriges lantbruksuniversitet
Carlos Alberto Arnillas, University of Toronto
Jonathan D. Bakker, College of the Environment
Elizabeth T. Borer, University of Minnesota Twin Cities
Kari Anne Bråthen, UiT Norges Arktiske Universitet
Marc W. Cadotte, University of Toronto
Clinton Carbutt, University of KwaZulu-Natal
Jane A. Catford, King's College London
Mary E. DuPre, MPG Ranch
Ciara Dwyer, Lunds Universitet
Nico Eisenhauer, German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig
Catalina Estrada, Imperial College London
Nicole Hagenah, University of Pretoria
Sylvia Haider, Leuphana Universität Lüneburg
Kyle E. Harms, Louisiana State University
Yann Hautier, Universiteit Utrecht
Erika I. Hersch-Green, Michigan Technological University
Johannes M.H. Knops, Xi'an Jiaotong-Liverpool University
Lauri Laanisto, Eesti Maaülikool
Ramesh Laungani, Marist University
Peter Macek, Eesti Maaülikool
Holly Martinson, McDaniel College
Jonathan Millett, Loughborough University
Meelis Pärtel, Ökoloogia ja Maateaduste Instituut
Steven C. Pennings, College of Natural Sciences and Mathematics
Pablo L. Peri, Instituto Nacional de Tecnología Agropecuaria Buenos Aires
Sally A. Power, Hawkesbury Institute for the Environment
Anita C. Risch, Eidgenössische Forschungsanstalt für Wald, Schnee und Landschaft WSL
Christiane Roscher, German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig
Eric W. Seabloom, University of Minnesota Twin Cities
Nicholas G. Smith, Texas Tech University

Document Type

Article

Publication Date

2-10-2026

Abstract

The search for predictors of plant diversity has challenged scientists for decades. Here we identify intense photosynthetically active radiation (PAR) as a major factor constraining plant species richness in global grasslands. We show that the strength of the negative relationship between species richness and PAR increases with increasing elevation and that species richness is more strongly correlated with intense PAR than with UV-B radiation, climate variables, and atmospheric nitrogen deposition. In addition to species richness, plant biomass was also negatively correlated with PAR at higher elevations, indicating that intense PAR also constrains plant biomass in montane grasslands. Furthermore, we show that the decrease in plant species richness with increasing PAR is mainly caused by a decrease in species richness of forbs, sedges, and rushes. In contrast, species richness of grasses was only negatively correlated with PAR at high elevations, and species richness of legumes was not significantly correlated with PAR. Our results suggest that PAR constrains plant species richness in global grasslands and limits the extent to which plant species of specific functional groups can migrate uphill in response to climate warming.

Publication Title

Proceedings of the National Academy of Sciences of the United States of America

Share

COinS