Contrasting pathways to tree longevity in gymnosperms and angiosperms

Authors

Roel J. Brienen, University of Leeds
Giuliano Maselli Locosselli, University of São Paulo
Stefan Krottenthaler, University of Passau
Emanuel Gloor, University of Leeds
Robyn Wrigley, University of Leeds
Steven L. Voelker, Michigan Technological University
Jan Altman, Institute of Botany of the Czech Academy of Sciences
Nela Altmanova, Institute of Botany of the Czech Academy of Sciences
Leander D. Anderegg, University of California Santa Barbara
Michele Baliva, Università della Tuscia
Deepak Barua, Indian Institute of Science Education and Research
Vaclav Bazant, Czech University of Life Sciences
Bryan Black, University of Arizona
Peter M Brown, Rocky Mountain Tree-Ring Research
Gregorio Ceccantini, University of São Paulo
R Justin DeRose, Utah State University
Jose Villanueva Diaz, Instituto Nacional de Investigaciones Forestales, Agricolas y Pecuarias
Alfredo Di Filippo, Università della Tuscia
Jiri Dolezal, Institute of Botany of the Czech Academy of Sciences
Louis Duchesne, Ministère des Ressources naturelles et des Forêts
Christopher Earle, Gymnosperm Database
Pavel Fibich, Institute of Botany of the Czech Academy of Sciences
Hardy Griesbauer, British Columbia Ministry of Forests
Samuli Helama, Natural Resources Institute Finland
Stefan Klesse, Swiss Federal Institute for Forest, Snow and Landscape Research
Kirill Korznikov, Institute of Botany of the Czech Academy of Sciences
David Lindenmayer, The Australian National University
Shuhui Liu, Beijing Forestry University
Lidio Lopez, Laboratorio de Dendrocronología e Historia Ambiental
Maurizio Mencuccini, Centre for Ecological Research and Forestry Applications (CREAF)
Thomas A. Nagel, University of Ljubljana

Document Type

Article

Publication Date

12-19-2025

Abstract

Tree longevity is thought to increase in growth-limiting, adverse environments, but a quantitative assessment of drivers of global variation in tree longevity is lacking. We assemble a global database of maximum longevity for 739 tree species and analyse associations between longevity and climate, soil, and species' functional traits. Our results show two primary pathways towards long lifespans. The first is slow growth in resource-limited environments, consistent with the "adversity begets longevity" paradigm. The second pathway is through relief from abiotic constraints in productive environments. Despite notable exceptions, long-lived gymnosperms tend to follow the first path through slow growth in cold environments, whereas long-lived angiosperms tend to follow the second ("productivity") path reaching maximum longevity generally in humid environments. For angiosperms, we identify two mechanisms for increased longevity under humid conditions. First, higher water availability increases species' maximum tree height which is associated with greater longevities. Secondly, greater water availability increases stand density and inter-tree competition, limiting growth which may increase tree lifespan. The documented differences between gymnosperm and angiosperm longevity are likely rooted in intrinsic differences in hydraulic architecture that provide fitness advantages for gymnosperms under high abiotic stress, and for angiosperms under increased productivity or competition.

Publication Title

Nature communications

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